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Why are box 1 wages (federal) and box 3 wages (Social Security) different?

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Why are box 1 wages (federal) and box 3 wages (Social Security) different?

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The following deductions reduce your federal, Social Security, and Medicare wages: out-of-pocket health expenses, van pool participation, DepCare, Health Flexible Spending Account, and parking. Back to top The following deductions reduce only federal and state grosses: 403(b) and Defined Contribution Plan (DC Plan) retirement.

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