What exactly do geneticists mean when they talk about “fitness”?
When geneticists or biologists speak of fitness they do not mean the kind of athletic conditioning that this word commonly brings to mind. Biological fitness is essentially the capability of a certain genotype to survive and reproduce successfully. Fitness measures the quantity of copies of genes (from a particular genotype) found in the next generation; it is a measure of reproductive success. Fitness can be and is measured and calculated by population geneticists.