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Does wheat grain when in an elevator consume oxygen or does it just produce toxic gases?

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Does wheat grain when in an elevator consume oxygen or does it just produce toxic gases?

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As a rare event, fermentation of grain within a grain elevator can decrease the oxygen concentration, creating the danger of asphyxiation for workers. See page 10 of: http://www.cdc.gov/niosh/pdfs/83-126c.

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