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Can Puritan tiger beetles exist in other habitats besides high cliffs?

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Can Puritan tiger beetles exist in other habitats besides high cliffs?

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All Maryland populations of the Puritan tiger beetle occur on high cliffs (over 30 feet in height) and on the beaches below and adjacent to them. Larval Puritan tiger beetles develop in borrows in the cliff face and in the breakdown material at the base of the cliff.

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